About 3 million years ago, the Isthmus of Panama (a narrow strip of land connecting North and South America) rose above sea level, dividing marine organisms into Pacific and Caribbean populations. Researchers examined species of snapping shrimp on both sides of the isthmus. Based on the morphospecies concept, there appeared to be seven pairs of species, with one species of each pair in the Pacific and the other in the Caribbean. Different species live at somewhat different depths in the ocean. Using mitochondrial DNA sequences, the researchers estimated phylogenies and found that these species pairs, separated by the isthmus, were indeed each other�s closest relatives. They investigated mating in the lab and found that many species pairs were not very interested in courting with each other, and any that did mate almost never produced fertile offspring. The sister populations on opposite sides of the isthmus are true species under which species concept? A. the morphospecies concept B. the biological species concept C. the phylogenetic species concept D. both A and C E. all of the above