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The chemiosmotic model requires an intact outer mitochondrial membrane. is supported by the observation that, when isolated mitochondria have an artificial electrochemical gradient (higher pH in the matrix) imposed on them, electrons spontaneously flow through the respiratory chain. is supported by the observation that isolated mitochondria that are actively respiring cause the pH of the solution they are in to increase. requires that mitochondrial ATP synthesis and electron flow through the respiratory chain be obligately coupled.
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