In Drosophila melanogaster, the wild-type eye colour is dark red. A fly from a true-breeding strain with scarlet eyes is crossed to a fly from a true-breeding strain with brown eyes. All the offspring have dark red eyes. The F1 flies are intercrossed to produce an F2 generation. This comprises 472 flies with dark-red eyes, 138 with scarlet eyes, 145 with brown eyes and 45 with white eyes. A. Three genes are segregating in this cross
B. Two independently assorting genes are segregating in this cross
C. Two linked genes are segregating in this cross
D. The phenotypes result from interactions between genes
E. The brown eye colour is the result of epistasis