1. Much of the behavior of all primates is learned. Nonetheless, we have suggested many times that primate behavior has been shaped by natural selection. How can natural selection shape behaviors that are learned

Nature selects best suited organisms having behaviours that help in the survival of organisms in that environment.

Explanation:

Natural selection can shaped the behaviors that are learned by the organisms because these behaviours helped organisms in their survival. Those organisms will be selected by the nature which can withstand the changing environmental conditions. Organisms adopt different behaviours in order to survive harsh environmental condition and these behaviour shaped by the natural selection in that organism so that they survive and grow in that environment.

just go with your heart buddy