Primary reinforcers are related to secondary reinforcers in which of the following ways?

You didn't provide us with any possible ways? Primary reinforcers are, however, types of reinforcers where they immediately elicit a reward or reward-like behavior once they are applied in a situation. An example would be apple juice. 

Secondary reinforcers on the other hand need to be associated with something for them to become reinforcers or they provide rewards in a non-direct way. An example would be money.

This is the main relationship between them.

Holozoic nutrition refers to the regular animal nutrition mode whereas saprophytic nutrition refers to the nutrition mode of fungi and bacteria.

Rna polymeras should attache to the promoter.