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He hypothesis that a mobile compound produced in the coleoptile tip controlled its bending toward directional light was tested by p. boysen-jensen in 1893. his experimental setup is represented below. both dark and light treatment groups had intact coleoptiles as controls. the experimental groups included: coleoptiles in which the tips were removed coleoptiles that were cut with the tips separated by mica coleoptiles that were cut with the tips separated by agar boysen-jensen left one set of coleoptiles in darkness for 24 hours and one set in directional light for 24 hours and observed whether they elongated and/or bended toward the light.
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